I think you.re quite right that there's no such thing as a pre-Trek Mantra.
I don't think the Mantra Pro counts as a prototype though, apparently there were over 200 made. My, far from original spec, one is very similar to your to pic, 1996, except for the long Fox air shock.
The later production Mantras have a far deeper flatter main frame (which doesn't catch your thighs as you pedal), the standard 1 1/8" headset on the Mantra comp and Fox shocks. The shocks on the later models are shorter, with a higher bottom mounting point. The main pivot is a nylon bearing (as seen on the 3rd and 4th pics above) this has an adjustable cone, with 2 pin holes in the pivot, the earlier Mantras have a non-adjustable, bronze bush, top 2 pics.
Some might say that Trek stopped Klein making all those small proprietary things that (depending on your point of view) made Kleins unique, or a pain in the ass to own. But the welding and paint finishes on the older Kleins are definitely better...
See OldKlein for some more technical development details.
http://www.oldklein.com/klein_evolution.htm
All the best,